Jellyfishes
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Anthozoa
Scyphozoa
Cubozoa

    The phylum Cnidaria ("ny-dare-ee-ah") includes an odd assortment of simple animals: sea anemones and jellyfishes. There is tissue level organization in these animals, but the only true organs are the gonads. They possess a nerve net that controls body activity, but there is no brain or centralized nervous control. The body is radially symmetrical. All species are aquatic or marine, though many, such as the anemones, can withstand many hours exposed to the air during low tides.

The phylum gets its name from the cnidocytes, specialized cells that contain hypodermic stinging structures attached to filaments, and small venom sacs. When these cells are in contact with prey, they fire the tiny hypodermic harpoon (called a nematocyst) and its long tube connecting with the venom sac. The microscopic size of the cnidocytes means that dozens to hundreds are typically fired simultaneously. The tubes keep prey attached to the tentacle until the venom induces paralysis. These microscopic dart-and-line structures are what give tentacles of jellyfishes and anemones their stick feeling when touched. Most cnidarians are harmless to humans. Some, such as the largest of them all, the lion's mane jellyfish, may cause severe pain. The Portuguese man-o'-war causes excruciating pain, but no human deaths can be attributed to the venom of this species. At the other extreme is the Australian sea wasp, the venom of which may cause death within minutes.

At left is a hydrozoan jellyfish. This Pacific resident possesses numerous stinging tentacles and a reddish luminescent region within the rim where the tentacles attach to the bell. This species grows to a total length of some 5 inches across the bell; the tentacles, when relaxed, may extend over 8 feet. The fresh water hydras, so often studied in biology classrooms, are probably the best known members of the class Hydrozoa. Photo by Dr. R. G. Sprackland.

The phylum contains four classes: the Hydrozoa (meaning "water animals"), which includes many jellyfishes, including several fresh-water species, and the hydras, which generally possess a velum; the Scyphozoa ("cup animals"), which includes the stinging sea nettle jellyfish; the Anthozoa ("flower animals"), which are the least like jellyfish in the phylum, including anemones, corals, and sea fans; and the Cubozoa ("box animals"), including box jellies and the most venomous animal known, the sea wasp jellyfish.

Jellyfishes (which are neither fishes nor closely related to vertebrates) have an unusual ability to shrink in size when food supplies are lacking, and growing again when food is plentiful.

Characteristics of Cnidaria include:

Radial body symmetry,
Two tissue levels (not three)--endoderm and ectoderm only,
A jelly like mesogleal layer between the endo- and ectodermal layers,
Cells with joint nutritive and muscular functions,
Epithelial muscular cells,
Cnidocytes with venom-conducting nematocysts,
Lack of any true organs excepting gonads,
Gastrovascular cavity present

Literature:  Click on a book to order a copy.

Mitchell, L., J. Mutchmor and W. Dolphin. 1988. Zoology. Benjamin/Cummings Publishing, Menlo Park, CA. ISBN: 0-8053-2562-X.

Rees, W.J. (editor). 1966. The Cnidaria and their Evolution. Symposia of the Zoological Society of London, No. 16. Academic Press, NY.

Sprackland, Robert. 1994. Jellyfish primer for the marine aquarist. Tropical Fish Hobbyist 42(9): 32-39.

White, Julian. 1995. CSL Antivenom Handbook. Commonwealth Serum Laboratories, Melbourne. ISBN: 0-646-26814-7.

Link:

Cnidaria WWW Server (jellyfishes & corals): www.ucihs.uci.edu/biochem/steele/default.html